V.L. Menable and A. Eritzan described in 1957 a new hexaploid species (2n=42) named T. zhukovskyi. This species occurs in Western Georgia in the same habitat as the tetraploid T. timopheevi. According to Jakubziner (Proc. 1st International Wheat Genetics Symposium, 1958, pages 207-220), T. zhukovshyi resembles T. timopheevi in ear morphology and growth characteristics and may have been derived from it. During a comparative study of the karyotypes of all hexaploid Triticum species, we found that T. zhukovskyi has a pair of satellited chromosomes possessing a supernumerary constriction of the type found in T. macha Dek. et Men. by Bhaduri and Ghosh (Cytologia 20, 148 -149, 1955). The secondary constriction and supernumerary constriction are separated from each other by a minute chromosome segment. The pair of Sat-chromosomes with supernumerary constriction present in T. zhukovskyi resembled one such pair in T. macha with regard to relative length and index value (short arm/long arm). While two such pairs occur in T. macha only one pair was found in T. zhukovskyi.

http://www.shigen.nig.ac.jp/wheat/wis/No13/p9-1/1.html